1. College of Earth Science and Engineering, Shandong University of Science and Technology, Qingdao, China
2. Key Laboratory of Evolutionary Systematics of Vertebrates, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, China
3. CAS Center for Excellence in Life and Paleoenvironment, Beijing, China
4. College of Palaeontology, Shenyang Normal University, Shenyang, China