1. Faculty of Sciences, Department of Geology, Mohamed I University, Oujda, Morocco
2. Histoire Naturelle de l’Homme Préhistorique (HNHP) - UMR7194, CNRS/Département Homme et Environnement/Muséum national d’Histoire naturelle/Sorbonne Université/UPVD, Département Homme et Environnement, Paris, France
3. SARChI Chair on Biodiversity Value and Change, Faculty of Science, Engineering and Agriculture, University of Venda, Thohoyandou, South Africa