1. School of the Earth Sciences and Resources, China University of Geosciences, Beijing, China
2. State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences, Beijing, China
3. Division of Vertebrate Paleontology, Carnegie Museum of Natural History, Pittsburgh, PA, USA
4. Mace Brown Museum of Natural History, Department of Geology and Environment Geosciences, College of Charleston, Charleston, South Carolina, USA
5. Dingshang Fossil Museum, Chengdu, Sichuan, China