1. a Colegio de la frontera Sur, Departamento de Ecologia Acuàtica , Chetumal, Quintana Roo, México
2. b Smithsonian Institution of Natural History , Washington, DC, USA
3. c Stazione Zoologica Anton Dohrn, Napoli, Group of Benthic Ecology (Ischia), Laboratory of Functional and Evolutionary Ecology , Napoli, Italy