1. aDivision of Bioscience and Bioindustry, Tokushima University, Tokushima, Japan;
2. Faculty of Life Science, Tokushima Bunri University, Tokushima, Japan;
3. Faculty of Medicine, Tokushima University, Tokushima, Japan;
4. Institute of Biomedical Sciences, Tokushima University Graduate School, Tokushima, Japan