1. Faculty of Sciences of Tunis, Biology Department, Research, Laboratory of Ecology, Biology and Physiology of Aquatic Organisms, University of Tunis El Manar, Tunis, Tunisia
2. Anatomy and Cytology Service, Mohamed Taher Maamouri Hospital, Nabeul, Tunisia
3. Aquatic Environment Exploitation Resources Unit, Higher Institute Fishing and Fish Farming of Bizerte, Menzel Jemil, Tunisia