1. UR: Physiology and Aquatic Environment, Department of Biology, Faculty of Sciences of Tunis, University Tunis El Manar, Tunis, Tunisia;
2. BVBGR-LR11ES31, Higher Institute of Biotechnology - Sidi Thabet, Biotechpole Sidi Thabet, University Manouba, Ariana, Tunisia;
3. Anatomy and Cytology Service, CHU Mohamed Taher Maamouri hospital, University Tunis El Manar, Tunis, Tunisia