1. School of Computational Science, Florida State University Tallahassee, Florida 32306, USA; E-mail: nylander@scs.fsu.edu
2. Department of Systematic Zoology, Evolutionary Biology Centre, Uppsala University Norbyvägen 18 D, SE-752 36 Uppsala, Sweden
3. Department of Zoology, Göteborg University Box 463, SE-405 30 Göteborg, Sweden
4. Department of Vertebrate Zoology and Molecular Systematics Laboratory, Swedish Museum of Natural History P.O. Box 50007, SE-104 05 Stockholm, Sweden
5. Current Address: Swedish Species Information Centre, Swedish University of Agricultural SciencesBox 7007, SE-750 07 Uppsala, Sweden
6. Department of Biodiversity and Conservation, Real Jardin Botanico-CSIC Plaza de Murillo 2; 28014 Madrid, Spain