1. Department of Herpetology, American Museum of Natural History Central Park West at 79th Street, New York, New York, 10024-5192, USA E-mail: rax@amnh.org (C.J.R.)
2. Division of Paleontology, American Museum of Natural History Central Park West at 79th Street, New York, New York, 10024-5192, USA
3. Département de Biologie Animale, Université d'Antananarivo BP 906, Antananarivo, 101, Madagascar
4. Center for Biodiversity and Conservation, American Museum of Natural History Central Park West at 79th Street, New York, New York, 10024-5192, USA