1. Department of Ecology and Evolutionary Biology, Yale University New Haven, Connecticut, USA
2. Current Address: Kewalo Marine Laboratory41 Ahui St., Honolulu, Hawaii, 96816, USA; E-mail: cdunn@hawaii.edu
3. National Oceanography Centre Southampton, SO14 3ZH, UK; E-mail: prp@noc.soton.ac.uk
4. Monterey Bay Aquarium Research Institute 7700 Sandholdt Road, Moss Landing, California, 95039-0628, USA; E-mail: haddock@mbari.org