1. School of Biological Sciences, Washington State University P.O. Box 644236, Pullman, Washington 99164-4236, USA; johnrobertclark@gmail.com (J.R.C.)
2. Center for Tropical Plant Science and Conservation, Marie Selby Botanical Gardens 811 S. Palm Avenue, Sarasota, Florida 34236, USA
3. Botany Department, Field Museum of Natural History 1400 S Lake Shore Drive, Chicago, Illinois, 60605, USA
4. Deparment of Botany, University of British Columbia Vancouver, BC V6T 1Z4, Canada
5. Department of Botany, National Museum of Natural History Washington, DC 20013, USA