1. Department of Ecology and Evolutionary Biology, University of Connecticut 75 N. Eagleville Road, Unit 3043, Storrs, Connecticut 06269–3043, USA E-mail: paul.lewis@uconn.edu (P.O.L.)
2. School of Computational Science and Information Technology (CSIT), 150-F Dirac Science Library, Florida State University Tallahassee, Florida 32306–4120, USA