1. Department of Plant Pathology
201F PSB, 1405 Veterans Drive, University of KentuckyLexington, KY 40546-0312, USA
2. Plant Biology Division, The Samuel Roberts Noble Foundation
2510 Sam Noble Parkway, Ardmore, OK 73401, USA
3. Department of Statistics, University of Kentucky
Lexington, KY 40526-0027, USA; E-mail: ruriko@ms.uky.edu (R.Y.) C.L.S., K.D.C., S.S. and R.Y. contributed equally to this work