1. Faculty of Science, Department of Biology, University of Mahasarakham, Tambon Khamriang Kantarawichai District 44150, Thailand; E-mail: claude@msu.ac.th(J.C.)
2. Chelonian Research Institute 401 South Central Avenue, Oviedo, Florida 32765, USA
3. 16, cours du Liégat, 75013 Paris, France
4. Institut des Sciences de l'Evolution, Université de Montpellier 2 UMR 5554 CNRS, Cc 064; 2, Place Eugène Bataillon, 34095 Montpellier Cedex 5, France