1. Department of Organismal and Evolutionary Biology, Harvard University 26 Oxford Street, Cambridge, Massachusetts 02138, USA; E-mail: kandul@fas.harvard.edu (N.P.K.)
2. Department of Entomology, St. Petersburg State University Universitetskaya nab. 7/9, St. Petersburg 199034, Russia
3. Center for Conservation Biology, Department of Biological Sciences, Stanford University Palo Alto California 94305, USA