1. Department of Systematic Zoology, Evolutionary Biology Centre, Uppsala University
Norbyvägen 18 D, SE-752 36, Uppsala, Sweden; E-mail: johan.nylander@ebc.uu.se (J.A.A.N)
2. Section of Ecology, Behavior and Evolution, Division of Biological Sciences, University of California–San Diego La Jolla
California 92093–0116, USA
3. Departamento de Bioversidad y Biología Evolutiva, Museo Nacional de Ciencias Naturales
José Gutiérrez Abascal 2, 28006 Madrid, Spain