1. Department of Ophthalmology and Visual Science, W.K. Kellogg Eye Center, University of Michigan, Ann Arbor, Michigan, USA
2. Department of Pathology, University of Michigan, Ann Arbor, Michigan, USA
3. Michigan Center for Translational Pathology, University of Michigan, Ann Arbor, Michigan, USA
4. Department of Human Genetics, University of Michigan, Ann Arbor, Michigan, USA
5. Rogel Cancer Center, University of Michigan, Ann Arbor, Michigan, USA