1. Laboratory of Ecology, Biology and Physiology of Aquatic organisms, Tunis Faculty of Science, University of Tunis El Manar, Tunis, Tunisia
2. University of Carthage, Higher Institute of Fisheries and Aquaculture of Bizerte (ISPAB), Bizerte, Tunisie
3. Station de recherche Sous-marines et Océanographiques (STARESO), Calvi, France
4. Laboratory of Oceanology, MARE Centre, University of Liège, Liege, Belgium