1. Department of Zoology and Entomology, University of the Free State, South Africa
2. Ongava Research Centre, Outjo, Namibia
3. Research Unit of Biodiversity, University of Oviedo, Mieres, Asturias, Spain
4. Mammal Research Institute, Department of Zoology and Entomology, University of Pretoria, South Africa
5. Department of Zoology, Stockholm University, Stockholm, Sweden