1. Department of Zoology, Stockholm University, Stockholm, Sweden
2. Department of Zoology, Swedish Museum of Natural History, Stockholm, Sweden
3. Department of Zoology and Entomology, University of the Free State, Phuthaditjhaba, South Africa
4. Research Unit of Biodiversity (UO-CSIC-PA), Spanish National Research Council, University of Oviedo, Mieres, Spain
5. Department of Zoology and Entomology, University of Pretoria, Pretoria, South Africa