1. Department of Biological Sciences, Higher Teacher’s Training College (ENS), University of Yaoundé I, Yaoundé, Cameroon
2. Department of Animal Biology and Physiology, University of Yaoundé I, Yaoundé, Cameroon
3. Institute of Evolutionary Science of Montpellier (ISEM), University of Montpellier (CNRS, IRD, EPHE), Montpellier, France