1. Department of Orthopaedics and Rehabilitation, Yale University School of Medicine, New Haven, CT, USA
2. Department of Molecular, Cell, and Developmental Biology, Yale University, New Haven, CT, USA
3. Department of Comparative Medicine and Molecular, Cellular, and Developmental Biology, Yale University School of Medicine, New Haven, CT, USA
4. Orentreich Foundation for the Advancement of Science, Cold Spring, NY, USA
5. The Center for Clinical and Translational Research, Maine Medical Center Research Institute, Scarborough, ME, USA