1. Department of Agricultural Sciences, Amar Telidji University, Laghouat, Algeria
2. Laboratory of Bio-Saharan Resources: Conservation and Valorization, Kasdi Merbah University, Ouargla, Algeria
3. Mediterranean Institute of Biodiversity and Ecology (IMBE), Avignon University, CNRS, IRD, Aix Marseille University, Avignon, France
4. Department of Biological Sciences, Kasdi Merbah University, Ouargla, Algeria
5. Laboratory Biodiversity and Conservation of Water and the Soils, University Abdelhamid Ibn Badis, Mostaganem, Algeria