1. Department of Plant Systematics, Bayreuth Center of Ecology and Environment Research (Bayceer), University of Bayreuth, Bayreuth, Germany
2. Department of Biological Sciences, Institute of Arts and Sciences, Far Eastern University, Manila, The Philippines
3. The Graduate School and Research Center for the Natural and Applied Sciences, University of Santo Tomas, Manila, The Philippines
4. The Philippine National Herbarium, National Museum of the Philippines, Manila, The Philippines