1. Muséum national d’Histoire naturelle, Institut de Systématique, Évolution, Biodiversité, UMR 7205 ISYEB MNHN/CNRS/UPMC/EPHE, Sorbonne Universités , Paris, France
2. State Natural History Museum NAS of Ukraine , Lviv, Ukraine
3. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences , Beijing, China
4. University of Chinese Academy of Sciences , Beijing, China