1. Senckenberg Centre for Human Evolution and Palaeoenvironment, University of Tübingen, Burgsteige 11, Tübingen, 72070, Germany
2. The Role of Culture in Early Expansions of Humans, University of Tübingen, Hölderlinstrasse 12, Tübingen, 72074, Germany
3. Department of Early Prehistory and Quaternary Ecology, University of Tübingen, Burgsteige 11, Tübingen, 72070, Germany
4. Evolutionary Studies Institute, University of the Witwatersrand, Braamfontein, 2000, Johannesburg, South Africa