1. Faculty of Sciences of Tunis, Department of Biology, University of Tunis El Manar, El-Manar II-Tunis, Tunisia
2. UMR CNRS 6027, IRDL, Univ. Bretagne Sud, Pontivy, France
3. Aridlands and Oases Cropping Laboratory, Institute of Arid Regions (IRA), Medenine, Tunisia
4. National Institute of Agronomic of Tunisia (INAT), Tunis, Tunisia