1. SeaWorld San Diego, San Diego, CA, USA;
2. Pacific Marine Mammal Center, Laguna Canyon Rd, Laguna Beach, CA, USA;
3. Department of Comparative, Diagnostic, and Population Medicine, College of Veterinary Medicine, University of Florida, Gainesville, FL, USA;
4. SeaWorld and Busch Gardens Species Preservation Laboratory, SeaWorld Parks and Entertainment, San Diego, CA, USA