1. Laboratory of Aromatic and Medicinal Plants, Biotechnology center of Borj-Cédria, BP 901, 2050 Hammam-lif, Tunisie
2. Department of Biology, College of Science, Hail University, P.O. Box 2440, Hail 2440, Saudi Arabia
3. Laboratory of Genetics, Biodiversity and Valorization of Bio-Resources (LR11ES41), Higher Institute of Biotechnology of Monastir, University of Monastir, Avenue Tahar Haddad, BP74, Monastir 5000, Tunisia
4. Department of Biology, College of Sciences, Taif University, P.O. Box 11099, Taif 21944, Saudi Arabia
5. Dipartimento di Farmacia Via Bonanno 6, 56126 Pisa, Italy