1. Breeding and Physiology of Medicinal Plants, Ferdowsi University of Mashhad, Iran
2. Department of Horticultural Science and Landscape Engineering, Faculty of Agriculture, Ferdowsi University of Mashhad, Iran
3. Department of Pharmacy, Faculty of Pharmacy, Mashhad University of Medical Sciences, Mashhad, Iran
4. Department of Biotechnology, Faculty of Agriculture, Ferdowsi University of Mashhad, Iran
5. Research Center for Plant Sciences, Ferdowsi University of Mashhad, Iran