1. Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, Japan
2. Field Science Center for Northern Biosphere, Hokkaido University, Hokkaido, Japan
3. Graduate School of Agriculture, Kyoto University, Kyoto, Japan
4. Field Science Education and Research Center, Kyoto University, Kyoto, Japan
5. Kyushu University Forest, Kyushu University, Fukuoka, Japan