1. Department of Forest and Biomaterials Science, Graduate School of Agriculture, Kyoto University, Ssakyo-ku, Kyoto, Japan
2. Hiraoka Forest Institute, Otsu, Japan
3. Faculty of Advanced Science and Technology, Ryukoku University, Otsu, Shiga Japan
4. Tohoku Research Center, Forestry and Forest Products Research Institute,Shimokuriyagawa, Morioka,Japan
5. Natural Resources Canada, Canadian Forest Service, Northern Forestry Centre, Edmonton, Canada