1. Laboratory of Forest Biology, Division of Forest and Biomaterials Science, Graduate School of Agriculture, Kyoto University, Kyoto, Japan
2. Forestry Technology Institute, Hyogo Prefectural Technology Center for Agriculture, Forestry and Fisheries, Hyogo, Japan
3. Department of Biology, Graduate School of Science, Kobe University, Hyogo, Japan
4. School of Human Science and Environment, University of Hyogo, Hyogo, Japan