1. Department of Entomology, University of Maryland, College Park, MD 20742, USA
2. Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013-7012, USA
3. Denver Museum of Nature & Science, Denver, CO 80205, USA
4. Department of Science Education, Gongju National University of Education, Woongjinro 27, Gongju, Chungnam 32553, Republic of Korea
5. BEES Program, University of Maryland, College Park, MD 20742, USA
6. College of Life Sciences, Capital Normal University, Beijing 100048, PR China