1. CAS Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla 666303, China;
2. University of Chinese Academy of Sciences, Beijing 100049, China;
3. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy Sciences, Beijing 100093, China;
4. Key Laboratory of Vertebrate Evolution and Human Origins, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China;
5. Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Beijing 100101, China;
6. Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650204, China