1. Department of Earth Sciences, University College London, Gower Street, London, WC1E 6BT, UK
2. Université de Lyon, Université Claude Bernard Lyon 1, CNRS, ENTPE, UMR 5023 LEHNA, F-69622, Villeurbanne, France
3. Department of Geosciences, University of Arizona, 1040 E 4th St., Tucson, AZ 85721, USA
4. Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, 10th St and Constitution Ave NW, Washington, DC 20013, USA
5. Division of Vertebrate Paleontology, Yale Peabody Museum of Natural History, 170 Whitney Ave, New Haven, CT 06520, USA
6. Department of Geology, Panjab University, Chandigarh, 160 014, India