1. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Palaeoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China
2. University of Chinese Academy of Sciences (UCAS), Beijing 100049, China
3. The Institute of Basic Science, Korea University, 145 Anam-ro, Seongbuk-gu, Seoul, 02841, Republic of Korea
4. Department of Life Sciences, Brunel University, Kingston Lane, Uxbridge, UB8 3PH, United Kingdom
5. Earth Sciences Department, Natural History Museum, Cromwell Road, London, SW7 5BD, UK