1. University of Colorado Museum of Natural History, University of Colorado, Boulder, CO 80309, USA
2. University of California Museum of Paleontology, University of California, Berkeley, CA 94720, USA
3. Department of Biological Sciences, Florida State University, Tallahassee, FL 32306, USA
4. University of Alaska Museum, University of Alaska, Fairbanks, AK 99775, USA
5. Department of Geosciences, University of Alaska Fairbanks, AK 99775, USA