1. Department of Biological Sciences, University of Alberta, Edmonton, Alberta T6G 2E9, Canada;
2. Department of Earth and Atmospheric Sciences, University of Alberta, Edmonton, Alberta T6G 2E9, Canada;
3. Department of Organismic and Evolutionary Biology & Museum of Comparative Zoology, Harvard University, Cambridge, MA 02138, USA;
4. College of Science and Engineering, Flinders University, Adelaide, SA 5042, Australia;
5. Earth Sciences Section, South Australian Museum, North Terrace, Adelaide, SA 5000, Australia;
6. Fundación de Historia Natural Félix de Azara, Centro de Ciencias Naturales Ambientales y Antropológicas, Universidad Maimónides, Hidalgo 775 (1405), Ciudad Autónoma de Buenos Aires, Argentina;
7. Department of Biology, University of Toronto Mississauga, Mississauga, Ontario L5L 1C6, Canada;
8. Arizona College of Osteopathic Medicine, Midwestern University, 19555 N. 59th Dr., Glendale, AZ 85383, USA