1. Department of Anatomy and Developmental Biology, Biomedicine Discovery Institute, Monash University, Melbourne, Victoria 3800, Australia
2. Vertebrate Palaeontology, Museums Victoria, Melbourne, Victoria 3001, Australia
3. School of Biological Sciences, Monash University, Melbourne, Victoria 3800, Australia
4. National Museum of Natural History, Smithsonian Institution, Washington DC 20560, USA
5. Department of Life Sciences, Natural History Museum, London SW7 5BD, UK