1. Department of Biology, College of Life and Environmental Sciences, Shanghai Normal University, Shanghai 200234, China;
2. Key Laboratory of Economic Stratigraphy and Palaeogeography, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China;
3. School of Earth Sciences, University of Bristol, Bristol BS8 1TQ, UK;
4. Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China;
5. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China;
6. Division of Entomology, Natural History Museum, and Department of Ecology & Evolutionary Biology, 1501 Crestline Drive – Suite 140, University of Kansas, Lawrence, Kansas 66045-4415, USA;
7. Division of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79th Street, New York, New York 10024-5192, USA