1. College of Life Sciences, Capital Normal University, Haidian District, Beijing 100048, China
2. Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washingto, DC, 20013-7012, USA
3. Department of Entomology and BEES Program, University of Maryland, College Park, MD, 20742, USA
4. Institute of Entomology, College of Plant Protection, Southwest Universitn, Beibei, Chongqing 400716, China
5. Northwest Institute of Plateau Biology, Chinese Academy of Sciences, 23, Xinning Road, 810008 Xining, China