1. Department of Biological Sciences, University of Alberta, Edmonton, Alberta T6G 2E9, Canada
2. Smithsonian Tropical Research Institute, Balboa–Ancón 0843–03092, Panama, Panama
3. Natural History Museum of Los Angeles County, 900 Exposition Blvd, Los Angeles, CA 90007, USA
4. Florida Museum of Natural History, PO Box 117800, University of Florida, Gainesville, FL 32611, USA
5. School of Life Sciences, PO Box 874501, Arizona State University, Tempe, AZ 85287-4501, USA