1. Key Laboratory of Vertebrate Evolution and Human Origins, of the Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, 100044, China;
2. CAS Center for Excellence in Life and Paleoenvironment, Beijing, 100044, China;
3. Kanagawa Prefectural Museum of Natural History, Odawara, Kanagawa, 250-0031, Japan;
4. Department of Earth’s Evolution and Environment, Graduate School of Science and Engineering, Ehime University, Ehime, 790-8577, Japan;
5. College of Earth and Planetary Sciences, University of Chinese Academy of Sciences, Beijing, 100049, China;
6. Department of Cell & Developmental Biology, University College London, Gower Street, WC1E 6BT, UK Biosciences, London