1. Zoological Institute of the Russian Academy of Sciences, Universitetskaya Emb. 1, 199034 Saint Petersburg, Russia;
2. Institute of Geology and Petroleum Technology, Kazan Federal University, Kremlevskaya Str. 4/5, 420008 Kazan, Russia;
3. Borissiak Paleontological Institute of the Russian Academy of Sciences, Profsouznaya Str. 123, 117647 Moscow, Russia