1. Department of Earth and Environmental Sciences and Museum of Paleontology, University of Michigan, Ann Arbor, MI 48103, USA
2. Department of Biological Sciences, University of Alberta, Edmonton, AB T6G 2E9, Canada
3. Department of Botany and Plant Pathology, 2082 Cordley Hall, Oregon State University, Corvallis, OR 97331, USA
4. Department of Environmental and Plant Biology, Ohio University, Athens, OH 45701, USA
5. Université Paris-Sud, Laboratoire Ecologie, Systématique, Evolution, CNRS UMR 8079, Orsay, 91405, France