1. Departament de Dinàmica de la Terra i de l'Oceà and Institut de Recerca de la Biodiversitat (IRBio), Facultat de Geologia, Universitat de Barcelona, Martí i Franquès s/n, Barcelona 08028, Spain
2. Division of Entomology, Natural History Museum, and Department of Ecology & Evolutionary Biology, University of Kansas, 1501 Crestline Drive, Lawrence, Kansas 66045-4415, USA
3. Division of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79th Street, New York, New York 10024-5192, USA