1. Institute of Geology and Paleontology, Linyi University, Shuangling Road, Linyi 276000, China;
2. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China;
3. Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1, Beichen West Road, Beijing 100101, China;
4. Department of Earth Sciences, Natural History Museum, Cromwell Road, London SW7 5BD, UK;
5. State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences, Xueyuan Lu 29, Beijing 100083, China