1. Department of Biology, College of Life and Environmental Sciences, Shanghai Normal University, Shanghai, 200234, China;
2. CAS Key Laboratory of Economic Stratigraphy and Palaeogeography, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China;
3. School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS8 1TQ, UK;
4. Center for Integrative Research, Field Museum of Natural History, Chicago, Illinois 60605, USA