1. Faculty of Natural Resources Management, Lakehead University, 955 Oliver Road, Thunder Bay, ON, P7B 5E1, Canada
2. Department of Geography and the Environment, Lakehead University, 955 Oliver Road, Thunder Bay, ON, P7B 5E1, Canada
3. Centre for Northern Forest Ecosystem Research, Ontario Ministry of Natural Resources and Forestry, 421 James St S, Thunder Bay, ON, P7E 2V6, Canada